Lanayru (Spirit)

Lanayru is one of the four Light Spirits in Twilight Princess, it protects the Lanayru Province and appears in the form of a serpent. Lanayru explains how the Fused Shadows, which Link is searching for, were formed by the power of a race of dark magic wielders. But the Gods summoned the four light spirits to intervene. The light spirits sealed away their magic into the fused shadows and the interlopers were chased across Hyrule into the Gerudo desert where they were then banished to the Twilight Realm by the Gods. Notice, the name "Lanayru", is a refrence to the goddess Nayru.

In the game the Lanayru spirit is the only one that has an intricate shrine built in its likeness around its spring, it is also mentioned that hylians make regular visits to pay respect to the spirit. When Zant attacks Midna and Link after finding the last fused shadow the Lanayru spirit attempts to attack him but is pushed back, this is the only time one of the spirits tries to attack an enemy directly.